Galium aparine

Family:RUBIACEAE
Species:Galium aparine L.
Common Name:Cleavers; Sticky Willy; Catchweed Bedstraw
Habitat:Various sites including woodlands, their edges, roadsides and other disturbed sites.
Associated Ecological Communities:**
Growth Habit:Vine
Duration:Annual
Category:Vascular
USDA Symbol:**
Plant Notes:Cleavers is a native or introduced herbaceous annual in the Madder family (Rubiaceae). Some authorities consider it native only to Europe, northern Africa, and Asia, and introduced in North America. It is possible that the species was introduced on the fur and clothing of people and animals crossing the Bering Land Bridge into North America from Asia during the last Ice Age as well as by colonists arriving from Europe in the 1600’s and 1700’s. In Alabama it can be found throughout the state. Cleavers occurs on roadsides, in forest edges and clearing, along streams, and in other disturbed sites. It is an annual with a tap root. The stems are prostrate or trailing over other vegetation and often form mats. The stems are green in color, angled, with short, stiff, backwards pointing teeth along their margins. The leaves occur in whorls of 6-8 per node. The leaves are sessile, linear to oblanceolate in outline, pubescent, with backwards pointing bristles along their margins. Flowers are produced in open, branched, terminal or axillary cymes. Individual flowers have 4 white petals. The fruit is a dry, globular nutlet with hooked bristles that occurs in pairs. Aparine is derived from Greek and means ‘clinging’ or ‘seizing’.—A. Diamond
Taxonomic Notes:**
Status:Native, FACU (NWI), G5 (Global Rank)
References:**

** Not applicable or data not available.

Classification

FamilyRUBIACEAE - Madder/Coffee family
Genus Galium
Species Galium aparine L. - Cleavers; Sticky Willy; Catchweed Bedstraw

Citation

Citation Galium aparine Linnaeus, Sp. Pl. 1: 108. 1753.
Basionym: **
Type: Without data (lectotype: LINN 129.3). Lectotypified by Jafri, in Jafri & El-Gadi, Fl. Libya 65: 15. 1979.

** Not applicable or data not available.

Synonyms

SynonymFull CitationBasionymType
Galium aparine var. intermedium Galium aparine Linnaeus, var. intermedium Bonnet, Petite Fl. Paris 191. 1883.Galium intermedium Merat 1831, non Schultes 1809. 
Galium aparine var. microphyllum Galium aparine Linnaeus, var. microphyllum Clos, in Gay, Fl. Chil. 3: 190. 1848. CHILE: Coquimbo: Without data, Gay 68 (holotype: P).
Galium aparine var. minor Galium aparine Linnaeus, var. minor Hooker, Fl. Bor.-Amer. 1: 290. 1834. CANADA: British Columbia: Near Fort Vancouver, s.d., Douglas s.n. (holotype: K?).
Galium aparine var. pseudoaparine Galium aparine Linnaeus, var. pseudoaparine (Grisebach) Spegazzini, Revista Agron. Univ. Nac. La Plata 3: 526. 1897.Galium pseudoaparine Grisebach 1854. 
Galium aparine var. subglabrum Galium aparine Linnaeus, var. subglabrum Petermann, Fl. Lips. Excurs. 122. 1838. GERMANY
Galium aparine var. verum Galium aparine Linnaeus, var. verum Wimmer & Grabowski, Fl. Siles. 1(1): 119. 1827, nom. inadmiss.Galium aparine Linnaeus 1753. 
Galium aparine var. vulgare Galium aparine Linnaeus, var. vulgare Reichenbach, Icon. Fl. Germ. Helv. 17: 94. 1854, nom. inadmiss.Galium aparine Linnaeus 1753. 
Galium australe Galium australe Reiche, Fl. Chile 3: 149. 1900, nom. illegit.Galium chilense Hooker f. 1846. 
Galium chilense Galium chilense Hooker f., Fl. Antarct. 302bis. 1846. CHILE: Asien: Chonos Archipelago, s.d., Darwin s.n. (holotype: K).
Galium chonosense Galium chonosense Clos, in Gay, Fl. Chil. 3: 189. 1848, nom. illegit. et non Hooker f. 1846.Galium chilense Hooker f. 1846. 
Galium hispidum Galium hispidum Willdenow, Enum. Pl. 154. 1809, non Roth 1789, nec Gaertner 1791. EGYPT
Galium intermedium Galium intermedium Merat, Nouv. Fl. Env. Paris, ed. 3. 2: 255. 1831, non Schultes 1809. FRANCE
Galium lappaceum Galium lappaceum Salisbury, Prodr. Stirp. Chap. Allerton 59. 1796, nom. illegit.Galium aparine Linnaeus 1753. 
Galium larecajense Galium larecajense Wernham, J. Bot. 50: 244. 1912. BOLIVIA: La Paz: Prov. Larecaja, vicinity of Sorata, between Munaypata and Challasuyo River, s.d., Mandon 339 (holotype: BM; isotype: NY).
Galium parviflorum Galium parviflorum Maximowicz, Prim. Fl. Amur. 479. 1859, non Moench 1794, nec Rafinesque 1808, nec D. Don 1825.  
Galium pauciflorum Galium pauciflorum Bunge, Enum. Pl. China Bor. 35. 1833, non Willdenow ex de Candolle 1830.  
Galium pseudoaparine Galium pseudoaparine Grisebach, Abh. Königl. Ges. Wiss. Göttingen 6: 125. 1854. CHILE: Magallanes: Near Punta Arenas, s.d., Lechler 1207 (holotype: GOET; isotypes: BR, G, K, P).
Galium scaberrimum Galium scaberrimum Vahl ex Hornemann, Hort. Bot. Hafn. 135. 1813.Galium hispidum Willdenow 1809, non Roth 1789, nec Gaertner 1791. 
Galium sororium Galium sororium Hance, in Walpers, Ann. Bot. Syst. 2: 734. 1852.  
Galium uncinatum Galium uncinatum Gray, Nat. Arr. Brit. Pl. 2: 484. 1821, nom. illegit.Galium aparine Linnaeus 1753. 
Galium aparine var. echinospermum    

Specimens and Distribution

This species has been reported in the following counties by the herbaria listed. An overview of the individual specimens are provided in the table that follows. Click on the accession number to view details; click on column headers to sort; choose a county or herbaria to filter the specimen data.

Counties included on distribution map: Autauga, Baldwin, Barbour, Blount, Bullock, Butler, Calhoun, Chambers, Cherokee, Chilton, Choctaw, Clarke, Clay, Cleburne, Coffee, Colbert, Conecuh, Coosa, Covington, Crenshaw, Cullman, Dale, Dallas, DeKalb, Elmore, Escambia, Etowah, Fayette, Geneva, Greene, Hale, Henry, Houston, Jackson, Jefferson, Lamar, Lauderdale, Lawrence, Lee, Limestone, Lowndes, Madison, Marengo, Marshall, Mobile, Monroe, Montgomery, Morgan, Pickens, Pike, Randolph, Russell, Shelby, St. Clair, Sumter, Tallapoosa, Tuscaloosa, Walker, Washington, Wilcox, Winston

Counties represented by specimen data listed below:

Herbaria represented by specimen data listed below: ALNHS, AUA, JSU, SAMF, TROY, UNA, USAM, UWAL, UWFP

Range of years during which specimens were collected: 1955 - 2000

Barcode / Accession No. County Coll. Date Collector &
Collection No.
Herbarium &
Herbarium Name Used
Image
UWAL0013444 Lawrence 27 Apr 2012 England, J. Kevin
3343
UWAL
Galium aparine Linnaeus
UNA00066662 Butler 22 Mar 2007 Diamond, Alvin R.
17331
UNA
Galium aparine L.
SAMF0004193 Montgomery 01 Apr 1965 Myers, Martha Crystal
s.n.
SAMF
Galium aparine Linnaeus
UWAL0037204 Cullman 08 Apr 2016 Spaulding, Daniel D.
14484
UWAL
Galium aparine L.
UNA00043517 Tuscaloosa 20 Apr 1955 Chermock, Ottilie D.
s.n.
UNA
Galium aparine L.
TROY000046251 Elmore 20 Mar 2018 Diamond, Alvin R.
28107
TROY
Galium aparine Linnaeus
AUA_ACC_41448 Cherokee 16 Apr 1972 Hale, A.R.
064
AUA
Galium Aparine L.
UWAL0049364 Bullock 29 Apr 2017 Horne, Howard E.
3826
UWAL
Galium aparine Linnaeus
JSU131854 Calhoun 20 Apr 1953 Pendergrass, Suevilla S.
s.n.
JSU
Galium aparine Linnaeus
TROY000022718 Marengo 06 Feb 2009 England, J. Kevin
1589
TROY
Galium aparine Linnaeus
JSU131937 Marshall 1 May 1973 Whetstone, R. David
RDW-1297
JSU
Galium aparine Linnaeus
TROY000023271 Geneva 03 Apr 2008 Diamond, Alvin R.
18835
TROY
Galium aparine Linnaeus
AUA_ACC_48679 Crenshaw 1 Apr 1988 Diamond, Alvin R.
4343
AUA
Galium Aparine L.
UWAL0046587 Autauga 04 Apr 2017 Parker, Matthew R.
544
UWAL
Galium aparine Linnaeus
AUA_ACC_14077 Calhoun 16 Apr 1968 O'Neal, Judy
AUA
Galium Aparine L.
TROY000043670 Limestone 25 Apr 2015 England, J. Kevin
5409
TROY
Galium aparine Linnaeus
JSU131850 Calhoun 16 Apr 1974 Atkinson, Tim A.
137
JSU
Galium aparine Linnaeus
JSU131934 Marshall 2 May 1992 Spaulding, Daniel D.
1503
JSU
Galium aparine Linnaeus
AUA_ACC_23814 Russell 3 Apr 1971 Donavan, Lois A.
053
AUA
Galium Aparine L.
UWAL0032305 Jackson 08 Apr 2016 England, J. Kevin
6095
UWAL
Galium aparine Linnaeus
UNA00054153 Blount 11 Apr 1998 Keener, Brian R.
581
UNA
Galium aparine L.
AUA_ACC_63885 Montgomery 22 Apr 2004 Diamond, Alvin R.
14347
AUA
Galium Aparine L.
JSU131931 Lee 8 Apr 1964 Bozarth, Gene A.
s.n.
JSU
Galium aparine Linnaeus
UNA00061301 Lawrence 20 Apr 2000 England, J. Kevin
133
UNA
Galium aparineáL.
TROY000053710 Barbour 24 Mar 2022 Diamond, Alvin R.
29838
TROY
Galium aparine Linnaeus
AUA_ACC_29208 Cleburne 14 May 1976 Rutland, David W.
2675
AUA
Galium Aparine L.
UWAL0050830 Greene 12 Apr 2021 Ryan, Emma D.
94
UWAL
Galium aparine L.
JSU131860 Colbert 24 May 1974 Whetstone, R. David
3190
JSU
Galium aparine Linnaeus
AUA_ACC_72616 Baldwin 1 Apr 2008 Barger, T. Wayne
PERD-315
AUA
Galium Aparine L.
UNA00043520 Dallas 09 Apr 1982 Gunn, Scott C.
396
UNA
Galium aparine L.
TROY000023272 Henry 13 Apr 2008 Diamond, Alvin R.
18976
TROY
Galium aparine Linnaeus
AUA_ACC_39349 Dallas 10 Apr 1982 Gunn, Scott C.
445
AUA
Galium Aparine L.
JSU131943 Walker 17 Apr 1980 Whetstone, R. David
9506
JSU
Galium aparine Linnaeus
UWAL0003896 Montgomery 30 Apr 2009 Diamond, Alvin R.
20644
UWAL
Galium aparine L.
TROY000008649 Coffee 03 Mar 2000 Martin, Brian H.
449
TROY
Galium aparine Linnaeus
AUA_ACC_45406 Chambers 14 Apr 1979 Carter, G.A.
033
AUA
Galium Aparine L.
JSU131924 Etowah 1 May 1992 Bailey, D.S.
66
JSU
Galium aparine Linnaeus
JSU131940 St. Clair 5 May 1983 Whetstone, R. David
13062
JSU
Galium aparine Linnaeus
JSU131929 Lauderdale 24 Apr 1969 Henson, Jim
s.n.
JSU
Galium aparine Linnaeus
TROY000008654 Montgomery 22 Apr 2004 Diamond, Alvin R.
14347
TROY
Galium aparine Linnaeus
AUA_ACC_9400 Lee 15 Apr 1960 Jones, Jr., Samuel B.
454
AUA
Galium Aparine L.
JSU131935 Marshall 13 Apr 1992 Spaulding, Daniel D.
1341
JSU
Galium aparine Linnaeus
UNA00043515 Dallas 11 Apr 1969 Thomas, Joab L.
1944
UNA
Galium aparine L.
AUA_ACC_65636 Lowndes 13 Apr 2006 Diamond, Alvin R.
16240
AUA
Galium Aparine L.
AUA_ACC_28931 Coosa 20 Apr 1976 Rutland, David W.
2286
AUA
Galium Aparine L.
UNA00044683 Randolph 18 Apr 1985 Nixon, C.F.
1084
UNA
Galium aparine L.
UNA00043527 Tuscaloosa 31 Dec 1957 Harper, Roland M.
4385
UNA
Galium aparine L.
UWAL0003894 Marengo 01 May 1993 Harris, J.W.
s.n.
UWAL
Galium aparine L.
UNA00043524 Tuscaloosa s.d. Smith, Eugene A.
s.n.
UNA
Galium aparine L.
TROY000008651 Dale 17 Mar 2000 Pennington, Tiffany
420
TROY
Galium aparine Linnaeus
UWAL0003892 Marengo 06 Feb 2009 England, J. Kevin
1589
UWAL
Galium aparine L.
AUA_ACC_48841 Monroe 2 Apr 1988 Diamond, Alvin R.
4369
AUA
Galium Aparine L.
JSU131859 Cleburne 25 Mar 1973 Windsor, Susan
s.n.
JSU
Galium aparine Linnaeus
TROY000023349 Lamar 06 Apr 2002 Spaulding, Daniel D.
11173
TROY
Galium aparine Linnaeus
AUA_ACC_61056 Choctaw 23 Apr 1994 Crouch, Virginia E.
882
AUA
Galium Aparine L.
JSU131858 Cleburne 22 Mar 1995 Hruska, Susan E.
576
JSU
Galium aparine Linnaeus
TROY000008650 Conecuh 07 Apr 2001 Diamond, Alvin R.
12348
TROY
Galium aparine Linnaeus
AUA_ACC_14619 Lee 8 May 1969 Barnes, W.K.
097
AUA
Galium Aparine L.
UWAL0003898 Sumter 10 Apr 2010 Keener, Brian R.
5751
UWAL
Galium aparine L.
TROY000023273 Houston 10 Apr 2008 Diamond, Alvin R.
18915
TROY
Galium aparine Linnaeus
AUA_ACC_10750 Lee 5 Apr 1963 Keyes, John W.
AUA
Galium Aparine L.
UWAL0039725 Elmore 20 Mar 2018 Diamond, Alvin R.
28107
UWAL
Galium aparine Linnaeus
TROY000023352 Tallapoosa 17 Mar 2002 Spaulding, Daniel D.
11116
TROY
Galium aparine Linnaeus
UWFP016190 Escambia 18 Apr 2010 James R. Burkhalter
22686
UWFP
Galium aparine L.
UNA00043519 Lee _ May 1897 Baker, Charles F.
s.n.
UNA
Galium aparine L.
UWAL0037344 Bullock 29 Mar 2016 Soehren, Eric C.
195
UWAL
Galium aparine L.
ALNHS00004393 Lee 04 Apr 2016 Taylor, Chris T.
CSP-289
ALNHS
Galium aparine L.
AUA_ACC_34763 Pike 30 Mar 1974 Jenkins, J.D.
053
AUA
Galium Aparine L.
JSU131925 Jackson 18 May 1999 Brodeur, Kristin R.
1819
JSU
Galium aparine Linnaeus
UNA00043522 Lawrence 09 May 1987 Haynes, Robert R.
9114
UNA
Galium aparine L.
AUA_ACC_29494 Cleburne 16 May 1976 Rutland, David W.
2763
AUA
Galium Aparine L.
UNA00043521 Tuscaloosa 27 Mar 1962 Gibbons, Whit
42
UNA
Galium aparine L.
UNA00043531 Hale 15 Apr 1978 Davenport, Lawrence J.
323
UNA
Galium aparine L.
AUA_ACC_41434 Elmore 17 Apr 1973 Harwell, W.Charles
146
AUA
Galium Aparine L.
UNA00043514 Hale 11 Apr 1966 Maginness, Sherrie E.
s.n.
UNA
Galium aparine L.
TROY000030539 Pike 06 Apr 1991 Mansfield, Shawn Lynn
2
TROY
Galium aparine Linnaeus
UWAL0003895 Morgan 08 Feb 2009 England, J. Kevin
1592
UWAL
Galium aparine L.
JSU131930 Jefferson 2 Apr 1976 Whetstone, R. David
s.n.
JSU
Galium aparine Linnaeus
USAM000013940 Mobile 14 Mar 1968 LeLong, Michel G.
4194
USAM
Galium aparine Linnaeus
JSU131921 DeKalb 20 Apr 1997 Ballard, J. Mark
7163
JSU
Galium aparine Linnaeus
TROY000023351 Pickens 06 Apr 2002 Spaulding, Daniel D.
11151
TROY
Galium aparine Linnaeus
AUA_ACC_35228 Butler 4 May 1979 Crawford, Lloyd C.
AUA
Galium Aparine L.
JSU131932 Madison 16 Apr 1996 Hruska, Susan E.
1030
JSU
Galium aparine Linnaeus
ALNHS00000742 Mobile 04 Apr 2010 Barger, T. Wayne
SP#275
ALNHS
Galium aparine L.
SAMF0004191 Cherokee 19 Apr 2010 Davenport, Lawrence J.
4238
SAMF
Galium aparine Linnaeus
AUA000037886 Covington 14 Mar 2012 Hansen, Curtis J.
5403
AUA
Galium Aparine L.
JSU131939 Randolph 18 Apr 1985 Nixon, Christopher F.
1084
JSU
Galium aparine Linnaeus
UWAL0018147 Limestone 25 Apr 2015 England, J. Kevin
5409
UWAL
Galium aparine Linnaeus
UNA00043523 Tuscaloosa 02 May 1959 Edwards, Jackie
s.n.
UNA
Galium aparine L.
AUA_ACC_49163 Crenshaw 22 Apr 1988 Diamond, Alvin R.
4516
AUA
Galium Aparine L.
UNA00043533 Marshall 23 Apr 1977 Ross, Larry J.
97
UNA
Galium aparine L.
UNA00065417 Winston 19 Apr 2005 England, J. Kevin
294
UNA
Galium aparine L.
TROY000035546 Bullock 22 Feb 2012 Dykes, Bailee
303
TROY
Galium aparine Linnaeus
AUA_ACC_27424 Randolph 13 Apr 1975 Rutland, David W.
272
AUA
Galium Aparine L.
UWAL0033653 Clarke 13 Mar 2016 Keener, Brian R.
9356
UWAL
Galium aparine Linnaeus
JSU131926 Jackson 20 May 1974 r, da
2713
JSU
Galium aparine Linnaeus
TROY000045773 Autauga 04 Apr 2017 Parker, Matthew R.
544
TROY
Galium aparine Linnaeus
JSU131851 Calhoun 11 May 1982 Whetstone, R. David
11025
JSU
Galium aparine Linnaeus
TROY000008653 Lowndes 13 Apr 2006 Diamond, Alvin R.
16240
TROY
Galium aparine Linnaeus
AUA_ACC_48713 Montgomery 10 Apr 1988 Diamond, Alvin R.
4400
AUA
Galium Aparine L.
JSU131938 Morgan 13 Apr 1997 Allison, James (Jim) R.
s.n.
JSU
Galium aparine Linnaeus
AUA000040208 Houston 22 Jan 2015 Barger, T. Wayne
DTP-12
AUA
Galium Aparine L.
TROY000049509 Lawrence 27 Apr 2012 England, J. Kevin
3343
TROY
Galium aparine Linnaeus
AUA_ACC_13038 Clay 17 Apr 1965 Brown, Jimmy E.
11
AUA
Galium Aparine L.
JSU131928 Jefferson 2 Apr 1976 Atkinson, Tim A.
1081
JSU
Galium aparine Linnaeus
JSU131942 Shelby 4 Apr 1993 Whetstone, R. David
16257
JSU
Galium aparine Linnaeus
UWAL0017645 Houston 02 Apr 2015 Diamond, Alvin R.
26087
UWAL
Galium aparine Linnaeus
UNA00062976 Montgomery 22 Apr 2004 Diamond, Alvin R.
14347
UNA
Galium aparine L.
UWAL0033881 Fayette 30 Mar 2016 Keener, Brian R.
9410
UWAL
Galium aparine Linnaeus
JSU131848 Blount 21 Apr 1973 Whetstone, R. David
RDW-1296
JSU
Galium aparine Linnaeus
TROY000008656 Pike 20 Mar 2003 Diamond, Alvin R.
13799
TROY
Galium aparine Linnaeus
AUA_ACC_45814 Houston 9 Apr 1971 Gwaltney, John R.
065
AUA
Galium Aparine L.
UWAL0009773 Dallas 04 Feb 2012 Keener, Brian R.
6840
UWAL
Galium aparine L.
JSU131849 Calhoun 22 Feb 1993 Dasinger, Adam P.
46
JSU
Galium aparine Linnaeus
TROY000023274 Wilcox 24 Mar 2008 Diamond, Alvin R.
18796
TROY
Galium aparine Linnaeus
JSU131933 Madison 1 May 1972 Whetstone, R. David
RDW-1268
JSU
Galium aparine Linnaeus
TROY000042454 Houston 02 Apr 2015 Diamond, Alvin R.
26087
TROY
Galium aparine Linnaeus
AUA_ACC_4357 Lee 25 Apr 1949 Cunningham, Hugh B.
AUA
Galium Aparine L.
UWAL0036100 Blount 19 Apr 2015 Anderson, Kelly
779
UWAL
Galium aparine L.
AUA_ACC_21930 Lauderdale 2 May 1970 Murchison, Grover R.
110
AUA
Galium Aparine L.
TROY000023269 Butler 27 Mar 2008 Diamond, Alvin R.
18824
TROY
Galium aparine Linnaeus
UWAL0003890 Covington 18 Mar 2008 Diamond, Alvin R.
18742
UWAL
Galium aparine L.
JSU131856 Clay 18 Mar 1995 Spaulding, Daniel D.
8903
JSU
Galium aparine Linnaeus
TROY000024641 Montgomery 30 Apr 2009 Diamond, Alvin R.
20644
TROY
Galium aparine Linnaeus
AUA_ACC_50401 Houston 9 Feb 1989 Diamond, Alvin R.
5676
AUA
Galium Aparine L.
UWAL0003897 St. Clair 07 Apr 2007 Keener, Brian R.
3445
UWAL
Galium aparine L.
TROY000044815 Geneva 23 Mar 2017 Byrd, Cameron
866
TROY
Galium aparine Linnaeus
AUA_ACC_41433 Washington 1 Apr 1973 Kelso, Emily
019
AUA
Galium Aparine L.
UWAL0003891 Jackson 12 Apr 2008 Keener, Brian R.
4222
UWAL
Galium aparine L.
TROY000008655 Pike 04 Apr 1997 Holmes, Jeanese
271
TROY
Galium aparine Linnaeus
UWAL0052032 Madison 11 Apr 2021 England, J. Kevin
11916
UWAL
Galium aparine L.
SAMF0008262 Walker 03 May 2017 Legg, Mary
43
SAMF
Galium aparine Linnaeus
AUA_ACC_34762 Pike 30 Mar 1974 Hutto, T.A.
107
AUA
Galium Aparine L.
JSU131855 Chilton 1 Apr 1999 Brodeur, Kristin R.
1597
JSU
Galium aparine Linnaeus
UNA00043513 Tuscaloosa 24 Feb 1955 Farmer, Joe A.
s.n.
UNA
Galium aparine L.
AUA_ACC_28738 Tallapoosa 15 Apr 1976 Rutland, David W.
2060
AUA
Galium Aparine L.
UNA00043525 Tuscaloosa 22 Feb 1957 Deason, Temd R.
s.n.
UNA
Galium aparine L.
JSU131922 Elmore 27 Apr 1979 Whetstone, R. David
9577
JSU
Galium aparine Linnaeus
AUA_ACC_48661 Pike 31 Mar 1988 Diamond, Alvin R.
4327A
AUA
Galium Aparine L.
UNA00048892 Henry 07 Apr 1997 MacDonald, John R.
10144
UNA
Galium aparine L.
TROY000023348 Chilton 14 Apr 2005 Diamond, Alvin R.
15140
TROY
Galium aparine Linnaeus
AUA_ACC_11389 Lee 9 Apr 1966 Searcy, James W.
AUA
Galium Aparine L.
JSU131941 Walker 17 Apr 1982 Whetstone, R. David
9492
JSU
Galium aparine Linnaeus
UWAL0043140 Baldwin 19 Mar 2000 Larsen, Harry S.
518
UWAL
Galium aparine L.
TROY000023350 Pickens 06 Apr 2002 Spaulding, Daniel D.
11141
TROY
Galium aparine Linnaeus
AUA_ACC_35216 Covington 27 Apr 1979 Coggins, W.A.
AUA
Galium Aparine L.
JSU131923 Etowah 6 May 1996 Hodge-Spaulding, Lesley M.
4458
JSU
Galium aparine Linnaeus
UNA00043518 Tuscaloosa 17 Apr 1963 Deramus, Rebecca
392
UNA
Galium aparine L.
SAMF0004192 Jefferson 21 Mar 2006 Davenport, Lawrence J.
3410
SAMF
Galium aparine Linnaeus
SAMF0004190 Tuscaloosa 15 Apr 1963 Williams, Norris H.
50
SAMF
Galium aparine Linnaeus
JSU131853 Calhoun 28 Mar 1994 Upton, D.L.
s.n.
JSU
Galium aparine Linnaeus
TROY000008648 Butler 22 Mar 2007 Diamond, Alvin R.
17331
TROY
Galium aparine Linnaeus
AUA_ACC_10676 Lee Taylor, T.D.
AUA
Galium Aparine L.
JSU131936 Marshall 13 Mar 1992 Spaulding, Daniel D.
1175
JSU
Galium aparine Linnaeus
UNA00043526 Tuscaloosa 17 Apr 1959 Williamson, Nelda Kathleen
s.n.
UNA
Galium aparine L.
AUA_ACC_39347 Dallas 9 Apr 1982 Gunn, Scott C.
396
AUA
Galium Aparine L.
AUA_ACC_29294 Tallapoosa 6 Apr 1976 Rutland, David W.
1996
AUA
Galium Aparine L.
JSU131852 Calhoun 18 Apr 1967 Collier, Gary
s.n.
JSU
Galium aparine Linnaeus
UNA00059122 Tallapoosa 17 Mar 2002 Spaulding, Daniel D.
11116
UNA
Galium aparine L.
JSU131927 Jefferson 30 Mar 1985 Barber, J. Patrick
1364
JSU
Galium aparine Linnaeus
UNA00059101 Pickens 06 Apr 2002 Spaulding, Daniel D.
11414
UNA
Galium aparine L.
TROY000008652 Dale 20 Feb 2000 Prazinko, Brian F.
s.n.
TROY
Galium aparine Linnaeus
UWAL0003893 Marengo 17 Apr 2009 England, J. Kevin
1941
UWAL
Galium aparine L.
AUA_ACC_20871 Colbert 23 Apr 1970 Morris, Janice
AUA
Galium Aparine L.
UNA00043516 Mobile 27 Mar 1965 Deramus, Rebecca
D286
UNA
Galium aparine L.
AUA_ACC_73097 Limestone 25 Apr 2015 England, J. Kevin
5409
AUA
Galium Aparine L.
JSU131857 Cleburne 26 Apr 1995 Hruska, Susan E.
668
JSU
Galium aparine Linnaeus
TROY000023270 Covington 18 Mar 2008 Diamond, Alvin R.
18742
TROY
Galium aparine Linnaeus
AUA_ACC_9706 Lee 5 May 1961 Beasley, P.G.
AUA
Galium Aparine L.
AUA_ACC_21931 Lee 17 Apr 1970 Williams, Ann
059
AUA
Galium Aparine L.
JSU131847 Baldwin 29 Mar 1998 Brodeur, Kristin R.
435
JSU
Galium aparine Linnaeus
UWAL0011456 Bullock 22 Feb 2012 Dykes, Bailee
303
UWAL
Galium aparine Linnaeus
UNA00060630 Pike 20 Mar 2003 Diamond, Alvin R.
13799
UNA
Galium aparine L.
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